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GRHL2 Entrez Description Chrom. Strand Promoter (Start - Stop) TSS 79977 grainyhead-like 2 (Drosophila) chr8 + 102499667 - 102505167 102504667 Transcription Factor Binding Sites   Download all TFBS in the GRHL2 promoter   Motif Source Strand Start Stop PValue Match Sequence Overlap w/ Footprints FOXO4_forkhead_DBD_dimeric_14_1 SELEX + 102506981 102506994 8.0E-06 AACAATGTGTTTAC 14 FOXJ2_forkhead_DBD_dimeric_13_1 SELEX + 102501317 102501329 1.0E-06 GAAAACAAATACA 13 FOXJ2_forkhead_DBD_dimeric_13_1 SELEX - 102505086 102505098 1.0E-06 AAAAACAAAAACA 13 SOX10_HMG_full_dimeric_16_1 SELEX - 102507484 102507499 2.0E-06 AAGAACATCCAGTGTG 16 IRX2_homeodomain_DBD_dimeric_12_1 SELEX - 102505079 102505090 6.0E-06 AAACATAACAAA 12 TBX5_TBX_DBD_dimeric_20_1 SELEX + 102500962 102500981 8.0E-06 GAGTGTGAAAGATTGCACCC 20 TBX5_TBX_DBD_dimeric_20_1 SELEX - 102500962 102500981 1.0E-05 GGGTGCAATCTTTCACACTC 20 NFYA_MA0060.1 JASPAR + 102504599 102504614 5.0E-06 CCCAGCCAATCATCGC 16 BARHL2_homeodomain_full_dimeric_16_1 SELEX - 102505392 102505407 8.0E-06 CAAATGGCCCTAAACG 16 NFATC1_NFAT_full_dimeric_14_1 SELEX - 102501137 102501150 7.0E-06 TTTTTCATGGGAAG 14 Foxc1_forkhead_DBD_dimeric_11_1 SELEX + 102501317 102501327 3.0E-06 GAAAACAAATA 11 FOXI1_MA0042.1 JASPAR - 102501319 102501330 3.0E-06 ATGTATTTGTTT 12 MGA_TBX_DBD_dimeric_18_2 SELEX + 102500963 102500980 1.0E-06 AGTGTGAAAGATTGCACC 18 MGA_TBX_DBD_dimeric_18_2 SELEX - 102500963 102500980 4.0E-06 GGTGCAATCTTTCACACT 18 TBX4_TBX_DBD_dimeric_20_1 SELEX + 102500962 102500981 6.0E-06 GAGTGTGAAAGATTGCACCC 20 FOXC2_forkhead_DBD_dimeric_11_1 SELEX + 102501317 102501327 4.0E-06 GAAAACAAATA 11 FOXC2_forkhead_DBD_dimeric_11_1 SELEX - 102505088 102505098 8.0E-06 AAAAACAAAAA 11 MZF1_5-13_MA0057.1 JASPAR - 102506223 102506232 4.0E-06 GGAGGGGGAA 10 ZSCAN4_C2H2_full_monomeric_15_1 SELEX + 102504923 102504937 9.0E-06 TGCACAGACTTGAAA 15 ZNF143_C2H2_DBD_monomeric_16_1 SELEX + 102504323 102504338 6.0E-06 GACCCACAATCCCTAG 16 TBX2_TBX_full_dimeric_18_1 SELEX + 102500963 102500980 3.0E-06 AGTGTGAAAGATTGCACC 18 TBX2_TBX_full_dimeric_18_1 SELEX - 102500963 102500980 3.0E-06 GGTGCAATCTTTCACACT 18 Tp53_p53l_DBD_dimeric_17_1 SELEX + 102505003 102505019 4.0E-06 ACAAGAGTCGGACAAGT 17 TBX20_TBX_DBD_monomeric_15_1 SELEX + 102500958 102500972 5.0E-06 GAAAGAGTGTGAAAG 15 FOXC1_forkhead_DBD_dimeric_13_1 SELEX + 102501315 102501327 2.0E-06 ACGAAAACAAATA 13 FOXC1_forkhead_DBD_dimeric_13_1 SELEX - 102505088 102505100 9.0E-06 TTAAAAACAAAAA 13 Foxj3_forkhead_DBD_dimeric_13_1 SELEX + 102501317 102501329 1.0E-06 GAAAACAAATACA 13 Foxj3_forkhead_DBD_dimeric_13_1 SELEX - 102505086 102505098 0.0E+00 AAAAACAAAAACA 13 FOXO1_forkhead_DBD_dimeric_14_1 SELEX + 102506981 102506994 5.0E-06 AACAATGTGTTTAC 14 IRX5_homeodomain_DBD_dimeric_12_1 SELEX - 102505079 102505090 3.0E-06 AAACATAACAAA 12 V_MEQ_01_M02049 TRANSFAC + 102507480 102507488 7.0E-06 AACACACAC 9 V_FOXP1_01_M00987 TRANSFAC + 102505079 102505098 0.0E+00 TTTGTTATGTTTTTGTTTTT 20 V_FOXP1_01_M00987 TRANSFAC + 102505085 102505104 1.0E-06 ATGTTTTTGTTTTTAAGTTG 20 V_CEBPG_Q6_M00622 TRANSFAC + 102501140 102501152 9.0E-06 CCCATGAAAAAAT 13 RXR_RAR_DR5_MA0159.1 JASPAR + 102506670 102506686 8.0E-06 GGGTCAGAGGCAGGGTA 17 V_IPF1_02_M01234 TRANSFAC - 102504304 102504313 2.0E-06 AGCTAATGAG 10 V_TCF3_01_M01594 TRANSFAC + 102505088 102505100 5.0E-06 TTTTTGTTTTTAA 13 V_AR_02_M00953 TRANSFAC - 102506973 102506999 1.0E-06 TGATGGTAAACACATTGTTCTTATTGC 27 V_FOXO4_02_M00476 TRANSFAC - 102501314 102501327 8.0E-06 TATTTGTTTTCGTC 14 V_AR_Q6_01_M01996 TRANSFAC - 102506978 102506992 1.0E-06 AAACACATTGTTCTT 15 V_EOMES_03_M02747 TRANSFAC + 102500958 102500974 5.0E-06 GAAAGAGTGTGAAAGAT 17 V_HNF4_Q6_M00967 TRANSFAC + 102504935 102504943 1.0E-05 AAAGTCCAG 9 V_FOXO3A_Q1_M01137 TRANSFAC + 102501316 102501327 8.0E-06 CGAAAACAAATA 12 V_FOXO3A_Q1_M01137 TRANSFAC - 102505088 102505099 1.0E-06 TAAAAACAAAAA 12 V_HNF3_Q6_01_M01012 TRANSFAC - 102501315 102501332 9.0E-06 TCATGTATTTGTTTTCGT 18 V_FOXJ1_03_M02750 TRANSFAC - 102506982 102506997 2.0E-06 ATGGTAAACACATTGT 16 V_STAT6_01_M00494 TRANSFAC - 102507463 102507470 7.0E-06 TATTTCCA 8 V_HFH4_01_M00742 TRANSFAC - 102501318 102501330 1.0E-06 ATGTATTTGTTTT 13 V_NFY_Q6_01_M00775 TRANSFAC + 102504598 102504610 6.0E-06 CCCCAGCCAATCA 13 V_CDX_Q5_M00991 TRANSFAC - 102505082 102505099 4.0E-06 TAAAAACAAAAACATAAC 18 V_CEBPB_01_M00109 TRANSFAC - 102507495 102507508 9.0E-06 GGATTATGAAAGAA 14 V_PAX_Q6_M00808 TRANSFAC + 102507462 102507472 1.0E-06 CTGGAAATAAC 11 V_AR_03_M00956 TRANSFAC - 102506973 102506999 1.0E-06 TGATGGTAAACACATTGTTCTTATTGC 27 V_CETS1P54_02_M00074 TRANSFAC + 102507537 102507549 9.0E-06 GCACAGGAAGATA 13 V_ELF4_04_M02850 TRANSFAC - 102505085 102505101 5.0E-06 CTTAAAAACAAAAACAT 17 V_HMBOX1_01_M01456 TRANSFAC + 102507548 102507564 3.0E-06 TATTTCTGGTTAACAAG 17 V_SOX13_03_M02797 TRANSFAC + 102506976 102506991 1.0E-06 ATAAGAACAATGTGTT 16 V_SOX7_03_M02807 TRANSFAC - 102505081 102505102 1.0E-06 ACTTAAAAACAAAAACATAACA 22 V_SOX7_03_M02807 TRANSFAC + 102506974 102506995 7.0E-06 CAATAAGAACAATGTGTTTACC 22 V_CRX_Q4_M00623 TRANSFAC + 102507499 102507511 4.0E-06 TTCATAATCCCAT 13 V_FOXA2_02_M02853 TRANSFAC - 102505076 102505090 3.0E-06 AAACATAACAAACCA 15 V_PR_Q2_M00960 TRANSFAC - 102507492 102507501 0.0E+00 GAAAGAACAT 10 V_TGIF_02_M01346 TRANSFAC - 102506547 102506563 2.0E-06 AAGTATGACAGCGCATT 17 V_PR_01_M00954 TRANSFAC - 102506973 102506999 1.0E-06 TGATGGTAAACACATTGTTCTTATTGC 27 V_ZFP105_03_M02827 TRANSFAC - 102505085 102505099 2.0E-06 TAAAAACAAAAACAT 15 V_ZFP105_03_M02827 TRANSFAC - 102505088 102505102 6.0E-06 ACTTAAAAACAAAAA 15 V_BEL1_B_M00312 TRANSFAC - 102500964 102500991 3.0E-06 TAACTGCTTAGGGTGCAATCTTTCACAC 28 V_FOXO1_02_M00474 TRANSFAC - 102501314 102501327 6.0E-06 TATTTGTTTTCGTC 14 V_STAF_01_M00262 TRANSFAC + 102504321 102504342 6.0E-06 TCGACCCACAATCCCTAGTGTT 22 Ddit3_Cebpa_MA0019.1 JASPAR - 102507531 102507542 7.0E-06 CTGTGCAATCCC 12 V_NFKB_Q6_M00194 TRANSFAC + 102506339 102506352 7.0E-06 AGGGGACCTTCCTG 14 V_FOXK1_04_M02856 TRANSFAC - 102505078 102505092 6.0E-06 AAAAACATAACAAAC 15 V_EVI1_01_M00078 TRANSFAC + 102501060 102501075 8.0E-06 AGAGATTATAAGATAT 16 V_HNF3_Q6_M00791 TRANSFAC - 102505086 102505098 9.0E-06 AAAAACAAAAACA 13 V_HNF3ALPHA_Q6_M00724 TRANSFAC - 102501317 102501327 8.0E-06 TATTTGTTTTC 11 V_CDX2_Q5_M00729 TRANSFAC - 102501063 102501076 8.0E-06 AATATCTTATAATC 14 V_GR_01_M00955 TRANSFAC - 102506973 102506999 4.0E-06 TGATGGTAAACACATTGTTCTTATTGC 27 V_IK2_01_M00087 TRANSFAC + 102507527 102507538 6.0E-06 CTTTGGGATTGC 12 V_GR_Q6_M00192 TRANSFAC - 102506977 102506995 3.0E-06 GGTAAACACATTGTTCTTA 19 V_NKX3A_01_M00451 TRANSFAC - 102506556 102506567 4.0E-06 TTTTAAGTATGA 12 V_STAT4_Q5_M02117 TRANSFAC + 102506749 102506758 5.0E-06 TTCAAGGAAA 10 V_TGIF1_01_M03111 TRANSFAC - 102506547 102506563 2.0E-06 AAGTATGACAGCGCATT 17 V_SOX11_03_M02795 TRANSFAC - 102505084 102505100 1.0E-06 TTAAAAACAAAAACATA 17 V_MZF1_02_M00084 TRANSFAC + 102504846 102504858 8.0E-06 TGGTGAGGGGGGA 13 V_MZF1_02_M00084 TRANSFAC - 102506223 102506235 1.0E-05 AGAGGAGGGGGAA 13 V_SOX9_B1_M00410 TRANSFAC + 102506977 102506990 4.0E-06 TAAGAACAATGTGT 14 V_ISGF4G_04_M02875 TRANSFAC - 102505080 102505093 3.0E-06 CAAAAACATAACAA 14 V_HNF4A_02_M02868 TRANSFAC + 102504931 102504946 5.0E-06 CTTGAAAGTCCAGTTT 16 V_FOXO1_Q5_M01216 TRANSFAC + 102501317 102501325 5.0E-06 GAAAACAAA 9 V_FOXO1_Q5_M01216 TRANSFAC - 102505090 102505098 1.0E-06 AAAAACAAA 9 V_GR_Q6_02_M01836 TRANSFAC - 102506977 102506989 4.0E-06 CACATTGTTCTTA 13 V_PTF1BETA_Q6_M00657 TRANSFAC + 102501083 102501096 3.0E-06 GAGAAACTGACAAG 14 V_CAAT_01_M00254 TRANSFAC + 102504599 102504610 3.0E-06 CCCAGCCAATCA 12 V_SOX15_03_M02799 TRANSFAC + 102506976 102506992 5.0E-06 ATAAGAACAATGTGTTT 17 V_CAAT_C_M00200 TRANSFAC + 102504603 102504627 2.0E-06 GCCAATCATCGCGCAGCTCGTCCCT 25 V_FOXL1_02_M02857 TRANSFAC - 102505076 102505091 0.0E+00 AAAACATAACAAACCA 16 V_FOXL1_02_M02857 TRANSFAC - 102505087 102505102 5.0E-06 ACTTAAAAACAAAAAC 16 V_FOXJ3_06_M02855 TRANSFAC - 102505076 102505092 2.0E-06 AAAAACATAACAAACCA 17 V_DMRT7_01_M01151 TRANSFAC + 102505080 102505093 5.0E-06 TTGTTATGTTTTTG 14 V_NFY_01_M00287 TRANSFAC + 102504599 102504614 4.0E-06 CCCAGCCAATCATCGC 16 V_HNF1B_01_M01425 TRANSFAC - 102507549 102507565 7.0E-06 ACTTGTTAACCAGAAAT 17 V_FOXO3_01_M00477 TRANSFAC - 102501314 102501327 4.0E-06 TATTTGTTTTCGTC 14 V_IRF6_04_M02874 TRANSFAC - 102504835 102504849 8.0E-06 ACCACTCTCGCTCGC 15 V_SMAD_Q6_01_M00974 TRANSFAC + 102501278 102501288 9.0E-06 TTTGCAGACAG 11 V_MEF3_B_M00319 TRANSFAC + 102501228 102501240 7.0E-06 TGGTGAGGTTTCT 13 V_RXR_RAR_01_M02272 TRANSFAC + 102506670 102506686 8.0E-06 GGGTCAGAGGCAGGGTA 17 V_DMRT2_01_M01147 TRANSFAC - 102504984 102504999 7.0E-06 ATGTTTGATCCAATGA 16 V_SRF_06_M02916 TRANSFAC - 102505084 102505100 1.0E-06 TTAAAAACAAAAACATA 17 V_SRF_06_M02916 TRANSFAC - 102505085 102505101 0.0E+00 CTTAAAAACAAAAACAT 17 V_SRF_06_M02916 TRANSFAC - 102505086 102505102 8.0E-06 ACTTAAAAACAAAAACA 17 V_HFH3_01_M00289 TRANSFAC - 102501318 102501330 2.0E-06 ATGTATTTGTTTT 13 V_FOXJ2_01_M00422 TRANSFAC + 102501315 102501332 2.0E-06 ACGAAAACAAATACATGA 18 V_FOXO1_01_M00473 TRANSFAC - 102505090 102505099 9.0E-06 TAAAAACAAA 10 V_SOX2_Q6_M01272 TRANSFAC + 102505084 102505099 1.0E-06 TATGTTTTTGTTTTTA 16 V_SOX2_Q6_M01272 TRANSFAC - 102506977 102506992 3.0E-06 AAACACATTGTTCTTA 16 V_PR_02_M00957 TRANSFAC - 102506973 102506999 2.0E-06 TGATGGTAAACACATTGTTCTTATTGC 27 V_SOX12_03_M02796 TRANSFAC + 102505088 102505101 4.0E-06 TTTTTGTTTTTAAG 14 V_SOX12_03_M02796 TRANSFAC - 102506975 102506988 2.0E-06 ACATTGTTCTTATT 14 V_SRY_05_M02917 TRANSFAC + 102506975 102506991 6.0E-06 AATAAGAACAATGTGTT 17 V_UF1H3BETA_Q6_M01068 TRANSFAC - 102506219 102506232 6.0E-06 GGAGGGGGAAGGGA 14 V_MAFK_04_M02880 TRANSFAC + 102506570 102506584 6.0E-06 CAAAAAACTGCCAGA 15 V_NFYA_Q5_M02106 TRANSFAC + 102504601 102504614 2.0E-06 CAGCCAATCATCGC 14 V_TCF3_05_M02920 TRANSFAC + 102501313 102501327 8.0E-06 GGACGAAAACAAATA 15 V_DEC_Q1_M00997 TRANSFAC + 102501035 102501047 8.0E-06 AGCCAAGTGAATG 13 V_DEC_Q1_M00997 TRANSFAC + 102504773 102504785 6.0E-06 CCCCATGTGAGGG 13 V_STAT1_05_M01260 TRANSFAC + 102507456 102507477 9.0E-06 TAGTTTCTGGAAATAACATGAA 22 V_FOXK1_03_M02752 TRANSFAC - 102506983 102506999 9.0E-06 TGATGGTAAACACATTG 17 V_PAX4_04_M00380 TRANSFAC - 102505062 102505091 3.0E-06 AAAACATAACAAACCAGTTGCCATCAGCCC 30 V_PAX4_04_M00380 TRANSFAC - 102505093 102505122 3.0E-06 GAAAAATACCAACAATAGCAACTTAAAAAC 30 V_SMAD1_01_M01590 TRANSFAC - 102505087 102505098 2.0E-06 AAAAACAAAAAC 12 V_NANOG_02_M01247 TRANSFAC - 102505083 102505102 2.0E-06 ACTTAAAAACAAAAACATAA 20 V_HSF1_Q6_01_M02017 TRANSFAC + 102507455 102507468 0.0E+00 GTAGTTTCTGGAAA 14 V_OTX3_01_M01403 TRANSFAC - 102507498 102507514 8.0E-06 TCAATGGGATTATGAAA 17

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