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GRHL1 Entrez Description Chrom. Strand Promoter (Start - Stop) TSS Motifs 29841 grainyhead-like 1 (Drosophila) chr2 + 10086791 - 10092291 10091791 GRHL1_CP2_DBD_dimeric_10_1 GRHL1_CP2_DBD_dimer-of-dimers_17_1 GRHL1_CP2_full_dimeric_12_1 Transcription Factor Binding Sites   Download all TFBS in the GRHL1 promoter   Motif Source Strand Start Stop PValue Match Sequence Overlap w/ Footprints FOXF2_MA0030.1 JASPAR + 10094384 10094397 5.0E-06 CTAAGATAAACAGT 14 FOXC2_forkhead_DBD_monomeric_12_1 SELEX - 10094426 10094437 2.0E-06 TAAATCAATAAA 12 ZNF75A_C2H2_DBD_monomeric_12_1 SELEX + 10094307 10094318 3.0E-06 GATTTTCCCACC 12 HINFP1_C2H2_full_dimeric_20_1 SELEX - 10091512 10091531 8.0E-06 GCGCACCCGCCAACCTCCGC 20 FOXC1_forkhead_DBD_monomeric_11_1 SELEX - 10094427 10094437 5.0E-06 TAAATCAATAA 11 NFYA_MA0060.1 JASPAR + 10091698 10091713 5.0E-06 GCCGACCAATCAGGAG 16 NFYA_MA0060.1 JASPAR + 10091762 10091777 2.0E-06 GCCAGCCAATGAGCGG 16 E2F7_E2F_DBD_dimeric_14_1 SELEX + 10094309 10094322 8.0E-06 TTTTCCCACCTATT 14 TBX1_TBX_DBD_dimeric_19_1 SELEX - 10094301 10094319 6.0E-06 AGGTGGGAAAATCTGGCTT 19 TEAD1_TEA_full_monomeric_10_1 SELEX + 10094452 10094461 1.0E-06 CACATTCCAT 10 ONECUT1_CUT_full_monomeric_14_1 SELEX - 10094426 10094439 1.0E-06 TGTAAATCAATAAA 14 CUX2_CUT_DBD_monomeric_10_1 SELEX - 10094427 10094436 8.0E-06 AAATCAATAA 10 GRHL1_CP2_DBD_dimer-of-dimers_17_1 SELEX + 10091917 10091933 9.0E-06 AACCCGAAAGTCCAGTT 17 GRHL1_CP2_DBD_dimer-of-dimers_17_1 SELEX - 10091917 10091933 1.0E-06 AACTGGACTTTCGGGTT 17 FOXJ3_forkhead_DBD_dimeric_14_1 SELEX + 10094389 10094402 3.0E-06 ATAAACAGTTAACT 14 SP1_MA0079.2 JASPAR + 10091641 10091650 7.0E-06 CCCCGCCCCC 10 ONECUT2_CUT_DBD_monomeric_14_1 SELEX - 10094426 10094439 1.0E-06 TGTAAATCAATAAA 14 ZIC3_C2H2_full_monomeric_15_1 SELEX - 10092006 10092020 1.0E-05 GCCCCCCCGCGGCGG 15 HNF1B_homeodomain_full_dimeric_13_1 SELEX - 10094426 10094438 7.0E-06 GTAAATCAATAAA 13 TEAD3_TEA_DBD_monomeric_8_1 SELEX + 10094453 10094460 1.0E-05 ACATTCCA 8 TEAD4_TEA_DBD_monomeric_10_1 SELEX + 10094452 10094461 1.0E-06 CACATTCCAT 10 PLAG1_MA0163.1 JASPAR - 10094441 10094454 4.0E-06 GTGGCCAAAGGGGA 14 FOXC1_forkhead_DBD_dimeric_13_1 SELEX - 10094424 10094436 2.0E-06 AAATCAATAAAGA 13 GLIS2_C2H2_DBD_monomeric_14_1 SELEX - 10092007 10092020 6.0E-06 GCCCCCCCGCGGCG 14 ONECUT3_CUT_DBD_monomeric_14_1 SELEX - 10094426 10094439 0.0E+00 TGTAAATCAATAAA 14 ZIC1_C2H2_full_monomeric_14_1 SELEX - 10092007 10092020 5.0E-06 GCCCCCCCGCGGCG 14 Zfx_MA0146.1 JASPAR - 10091736 10091749 6.0E-06 GGGGGCGAGGCCTC 14 FOXJ2_forkhead_DBD_dimeric_14_1 SELEX + 10094389 10094402 3.0E-06 ATAAACAGTTAACT 14 V_HSF2_01_M00147 TRANSFAC + 10094484 10094493 1.0E-06 AGAATATTCT 10 V_HSF2_01_M00147 TRANSFAC - 10094484 10094493 1.0E-06 AGAATATTCT 10 V_HNF3B_01_M00131 TRANSFAC + 10090808 10090822 1.0E-05 TTACTTGTTTTTGTA 15 V_HNF3B_01_M00131 TRANSFAC - 10094386 10094400 7.0E-06 TTAACTGTTTATCTT 15 V_DBX1_01_M01483 TRANSFAC - 10094424 10094440 6.0E-06 CTGTAAATCAATAAAGA 17 V_FREAC7_01_M00293 TRANSFAC - 10094371 10094386 4.0E-06 TAGAGATAAACACTTA 16 V_AHRARNT_01_M00235 TRANSFAC + 10094512 10094527 3.0E-06 GGAGGATTGCGTGAAG 16 V_NF1_Q6_01_M00806 TRANSFAC - 10094445 10094461 9.0E-06 ATGGAATGTGGCCAAAG 17 V_ZFX_01_M01593 TRANSFAC - 10091731 10091746 3.0E-06 GGCGAGGCCTCGGGGC 16 V_ZFX_01_M01593 TRANSFAC + 10091733 10091748 4.0E-06 CCCGAGGCCTCGCCCC 16 V_MTF1_Q4_M00650 TRANSFAC - 10092050 10092063 5.0E-06 TCTGCGCCCGGCTC 14 V_BCL6_01_M01183 TRANSFAC - 10094288 10094303 0.0E+00 CTTTTAAAAAATCTTT 16 V_BCL6_01_M01183 TRANSFAC + 10094321 10094336 0.0E+00 TTTTTTTTAAAACTTT 16 V_BCL6_01_M01183 TRANSFAC + 10094323 10094338 1.0E-06 TTTTTTAAAACTTTTT 16 V_MMEF2_Q6_M00405 TRANSFAC - 10094319 10094334 3.0E-06 AGTTTTAAAAAAAATA 16 V_SP1_Q6_01_M00931 TRANSFAC - 10091676 10091685 4.0E-06 GGGGCGGGGC 10 V_HNF4_Q6_M00967 TRANSFAC + 10091923 10091931 1.0E-05 AAAGTCCAG 9 V_NKX25_Q6_M02108 TRANSFAC - 10089795 10089805 5.0E-06 CTCACTTGAGA 11 V_CEBP_Q2_M00190 TRANSFAC - 10094263 10094276 2.0E-06 ACATTGCACAAGAA 14 V_HNF3_Q6_01_M01012 TRANSFAC - 10094383 10094400 3.0E-06 TTAACTGTTTATCTTAGA 18 V_SP1_03_M02281 TRANSFAC + 10091641 10091650 7.0E-06 CCCCGCCCCC 10 V_NERF_Q2_M00531 TRANSFAC - 10093140 10093157 5.0E-06 AGTCAGGAAGTAGGCTTC 18 V_AREB6_04_M00415 TRANSFAC - 10094254 10094262 8.0E-06 CTGTTTCAT 9 V_AREB6_04_M00415 TRANSFAC - 10095292 10095300 8.0E-06 CTGTTTCTA 9 V_SP1_02_M01303 TRANSFAC - 10091639 10091649 8.0E-06 GGGGCGGGGGC 11 V_SP1_02_M01303 TRANSFAC - 10091646 10091656 3.0E-06 GGGGCGGGGGG 11 V_CDX_Q5_M00991 TRANSFAC - 10090807 10090824 2.0E-06 AATACAAAAACAAGTAAA 18 V_RHOX11_02_M01384 TRANSFAC - 10096042 10096058 6.0E-06 AGTCTGCTGTTTTAAGT 17 V_TAL1_GATA1_01_M02243 TRANSFAC - 10094377 10094394 8.0E-06 GTTTATCTTAGAGATAAA 18 V_HNF1_C_M00206 TRANSFAC + 10094473 10094489 9.0E-06 TGGTAATCAGTAGAATA 17 V_NFY_C_M00209 TRANSFAC - 10091698 10091711 3.0E-06 CCTGATTGGTCGGC 14 V_ELF4_04_M02850 TRANSFAC - 10094288 10094304 1.0E-06 GCTTTTAAAAAATCTTT 17 V_ELF4_04_M02850 TRANSFAC - 10094316 10094332 1.0E-06 TTTTAAAAAAAATAGGT 17 V_ELF4_04_M02850 TRANSFAC - 10094317 10094333 8.0E-06 GTTTTAAAAAAAATAGG 17 V_ZNF219_01_M01122 TRANSFAC + 10091644 10091655 5.0E-06 CGCCCCCCGCCC 12 V_SPI1_03_M02078 TRANSFAC + 10093162 10093171 4.0E-06 TGAGGAAGTG 10 V_ZFP105_03_M02827 TRANSFAC - 10094317 10094331 9.0E-06 TTTAAAAAAAATAGG 15 V_ZFP105_03_M02827 TRANSFAC - 10094318 10094332 3.0E-06 TTTTAAAAAAAATAG 15 V_TBX18_01_M01262 TRANSFAC - 10094301 10094319 5.0E-06 AGGTGGGAAAATCTGGCTT 19 V_NKX31_02_M02782 TRANSFAC - 10094366 10094382 2.0E-06 GATAAACACTTAAGGAT 17 V_ZBP89_Q4_M01816 TRANSFAC + 10091888 10091897 6.0E-06 TCCTCCCCCC 10 V_HSF1_01_M00146 TRANSFAC + 10094484 10094493 3.0E-06 AGAATATTCT 10 V_HSF1_01_M00146 TRANSFAC - 10094484 10094493 3.0E-06 AGAATATTCT 10 V_SP1_Q6_M00196 TRANSFAC - 10091639 10091651 4.0E-06 GGGGGGCGGGGGC 13 V_EGR1_Q6_M01873 TRANSFAC + 10091459 10091468 4.0E-06 GCGGGGGCGG 10 V_SOX_Q6_M01014 TRANSFAC + 10092158 10092170 2.0E-06 CTCTTTGTTCGGT 13 V_LEF1_03_M02878 TRANSFAC - 10094423 10094438 9.0E-06 GTAAATCAATAAAGAG 16 V_NKX22_02_M01372 TRANSFAC - 10094365 10094381 3.0E-06 ATAAACACTTAAGGATA 17 V_PBX1_01_M00096 TRANSFAC - 10094426 10094434 5.0E-06 ATCAATAAA 9 V_NFE4_Q5_M02105 TRANSFAC + 10091547 10091558 2.0E-06 CTCCCTCTCCTG 12 Tal1_Gata1_MA0140.1 JASPAR - 10094377 10094394 8.0E-06 GTTTATCTTAGAGATAAA 18 V_SP1SP3_Q4_M01219 TRANSFAC + 10091883 10091893 5.0E-06 CCGCCTCCTCC 11 V_HMX1_02_M01481 TRANSFAC - 10094365 10094381 8.0E-06 ATAAACACTTAAGGATA 17 V_MEF2_04_M00233 TRANSFAC + 10096036 10096057 6.0E-06 GGTGCTACTTAAAACAGCAGAC 22 V_TEL2_Q6_M00678 TRANSFAC + 10093145 10093154 1.0E-06 CTACTTCCTG 10 V_CDC5_01_M00478 TRANSFAC - 10094322 10094333 8.0E-06 GTTTTAAAAAAA 12 V_ARID3A_04_M02735 TRANSFAC - 10094318 10094334 7.0E-06 AGTTTTAAAAAAAATAG 17 V_ARID3A_04_M02735 TRANSFAC + 10094319 10094335 9.0E-06 TATTTTTTTTAAAACTT 17 V_POU6F1_01_M00465 TRANSFAC + 10094347 10094357 8.0E-06 GCTTCATTTAT 11 V_TBP_06_M02814 TRANSFAC + 10094320 10094335 2.0E-06 ATTTTTTTTAAAACTT 16 V_FOXO1_Q5_M01216 TRANSFAC - 10090811 10090819 8.0E-06 AAAAACAAG 9 V_NKX32_02_M01482 TRANSFAC - 10094366 10094382 0.0E+00 GATAAACACTTAAGGAT 17 V_SP4_Q5_M01273 TRANSFAC + 10091640 10091650 5.0E-06 CCCCCGCCCCC 11 V_SP4_Q5_M01273 TRANSFAC + 10091647 10091657 7.0E-06 CCCCCGCCCCG 11 V_SP4_Q5_M01273 TRANSFAC + 10091676 10091686 2.0E-06 GCCCCGCCCCG 11 V_AML2_Q3_01_M01854 TRANSFAC + 10090799 10090809 1.0E-06 CCACACCATTT 11 V_ALPHACP1_01_M00687 TRANSFAC + 10091764 10091774 0.0E+00 CAGCCAATGAG 11 V_MTF1_06_M02882 TRANSFAC - 10094321 10094334 8.0E-06 AGTTTTAAAAAAAA 14 V_MTF1_06_M02882 TRANSFAC - 10094322 10094335 9.0E-06 AAGTTTTAAAAAAA 14 V_NFYC_Q5_M02107 TRANSFAC + 10091764 10091777 3.0E-06 CAGCCAATGAGCGG 14 V_CEBPA_01_M00116 TRANSFAC - 10094263 10094276 4.0E-06 ACATTGCACAAGAA 14 V_HOXB8_01_M01451 TRANSFAC - 10094349 10094364 8.0E-06 TCCAACAATAAATGAA 16 V_OSR2_04_M02889 TRANSFAC + 10096035 10096050 5.0E-06 GGGTGCTACTTAAAAC 16 V_RHOX11_05_M03099 TRANSFAC - 10096042 10096058 6.0E-06 AGTCTGCTGTTTTAAGT 17 V_NFY_01_M00287 TRANSFAC + 10091698 10091713 4.0E-06 GCCGACCAATCAGGAG 16 V_NFY_01_M00287 TRANSFAC + 10091762 10091777 2.0E-06 GCCAGCCAATGAGCGG 16 V_NKX22_01_M00485 TRANSFAC + 10094370 10094379 2.0E-06 TTAAGTGTTT 10 V_FPM315_01_M01587 TRANSFAC - 10091885 10091896 2.0E-06 GGGGGAGGAGGC 12 V_NRF2_Q4_M00821 TRANSFAC - 10091962 10091974 9.0E-06 CTCCTGTGTCATC 13 V_FOXO3_01_M00477 TRANSFAC + 10090809 10090822 5.0E-06 TACTTGTTTTTGTA 14 V_SPIC_02_M02077 TRANSFAC + 10093162 10093171 5.0E-06 TGAGGAAGTG 10 V_FOX_Q2_M00809 TRANSFAC + 10094425 10094437 2.0E-06 CTTTATTGATTTA 13 V_BBX_03_M02739 TRANSFAC - 10094423 10094437 4.0E-06 TAAATCAATAAAGAG 15 V_OSR1_04_M02888 TRANSFAC + 10096035 10096050 9.0E-06 GGGTGCTACTTAAAAC 16 V_SOX8_03_M02808 TRANSFAC + 10094350 10094366 4.0E-06 TCATTTATTGTTGGATA 17 V_SOX14_04_M02901 TRANSFAC - 10094308 10094324 4.0E-06 AAAATAGGTGGGAAAAT 17 V_SRF_06_M02916 TRANSFAC - 10094315 10094331 5.0E-06 TTTAAAAAAAATAGGTG 17 V_SRF_06_M02916 TRANSFAC - 10094316 10094332 1.0E-06 TTTTAAAAAAAATAGGT 17 V_SRF_06_M02916 TRANSFAC - 10094317 10094333 4.0E-06 GTTTTAAAAAAAATAGG 17 V_FOXJ2_01_M00422 TRANSFAC - 10094370 10094387 1.0E-06 TTAGAGATAAACACTTAA 18 V_FOXJ2_01_M00422 TRANSFAC + 10094383 10094400 0.0E+00 TCTAAGATAAACAGTTAA 18 V_FOXO1_01_M00473 TRANSFAC - 10090811 10090820 9.0E-06 CAAAAACAAG 10 V_SRY_05_M02917 TRANSFAC - 10094351 10094367 2.0E-06 ATATCCAACAATAAATG 17 V_FREAC4_01_M00292 TRANSFAC + 10094384 10094399 3.0E-06 CTAAGATAAACAGTTA 16 V_SP1_Q2_01_M00933 TRANSFAC + 10091641 10091650 4.0E-06 CCCCGCCCCC 10 V_FOXO4_01_M00472 TRANSFAC - 10090809 10090819 5.0E-06 AAAAACAAGTA 11 V_CEBP_Q2_01_M00912 TRANSFAC - 10094263 10094274 9.0E-06 ATTGCACAAGAA 12 V_LPOLYA_B_M00318 TRANSFAC - 10094425 10094432 7.0E-06 CAATAAAG 8 V_NFYA_Q5_M02106 TRANSFAC + 10091700 10091713 2.0E-06 CGACCAATCAGGAG 14 V_NFYA_Q5_M02106 TRANSFAC + 10091764 10091777 0.0E+00 CAGCCAATGAGCGG 14 V_SP1_Q4_01_M00932 TRANSFAC - 10091639 10091651 8.0E-06 GGGGGGCGGGGGC 13 V_PBX1_04_M01357 TRANSFAC - 10094424 10094440 5.0E-06 CTGTAAATCAATAAAGA 17 V_GLIS2_04_M02863 TRANSFAC - 10094321 10094334 8.0E-06 AGTTTTAAAAAAAA 14 V_AR_Q2_M00447 TRANSFAC + 10094329 10094343 1.0E-06 AAAACTTTTTGTTCA 15 V_PAX4_04_M00380 TRANSFAC + 10094412 10094441 3.0E-06 GAAAATTATGCCTCTTTATTGATTTACAGT 30 V_DMRT5_01_M01150 TRANSFAC - 10094250 10094264 1.0E-05 AACTGTTTCATTTTA 15 V_SPIB_03_M02076 TRANSFAC + 10093162 10093171 5.0E-06 TGAGGAAGTG 10 V_CEBPA_Q6_M01866 TRANSFAC + 10094263 10094275 7.0E-06 TTCTTGTGCAATG 13 V_HSF1_Q6_01_M02017 TRANSFAC - 10094406 10094419 4.0E-06 TAATTTTCTGGCAT 14

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