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CLUL1 Entrez Description Chrom. Strand Promoter (Start - Stop) TSS 27098 clusterin-like 1 (retinal) chr18 + 591997 - 597497 596997 Transcription Factor Binding Sites   Download all TFBS in the CLUL1 promoter   Motif Source Strand Start Stop PValue Match Sequence Overlap w/ Footprints CTCF_MA0139.1 JASPAR + 597086 597104 2.0E-06 ACGCCTCCAGGGGGCAGCA 19 HOXD12_homeodomain_DBD_monomeric_9_1 SELEX + 598203 598211 2.0E-06 CTAATAAAA 9 Hoxc10_homeodomain_DBD_monomeric_10_2 SELEX + 598203 598212 3.0E-06 CTAATAAAAA 10 THRB_nuclearreceptor_DBD_dimeric_20_1 SELEX + 598308 598327 7.0E-06 GTGACCTTGAAGAAAGTCAC 20 THRB_nuclearreceptor_DBD_dimeric_20_1 SELEX - 598308 598327 3.0E-06 GTGACTTTCTTCAAGGTCAC 20 SOX4_HMG_DBD_dimeric_16_1 SELEX + 597259 597274 8.0E-06 GAACAAAGCCACTGTT 16 SOX4_HMG_DBD_dimeric_16_1 SELEX - 597260 597275 4.0E-06 GAACAGTGGCTTTGTT 16 SOX10_HMG_full_dimeric_16_1 SELEX - 596659 596674 3.0E-06 AAGAACATTCAGTTTT 16 MSX2_homeodomain_DBD_dimeric_18_1 SELEX + 598203 598220 6.0E-06 CTAATAAAAATGCCATAA 18 SOX10_HMG_full_dimeric_14_1 SELEX + 596664 596677 6.0E-06 TGAATGTTCTTTCC 14 ESRRA_nuclearreceptor_DBD_dimeric_19_1 SELEX - 598298 598316 3.0E-06 CAAGGTCACGCAAAGTTTA 19 SRY_HMG_DBD_dimeric_15_1 SELEX + 597260 597274 4.0E-06 AACAAAGCCACTGTT 15 SRY_HMG_DBD_dimeric_15_1 SELEX - 597260 597274 7.0E-06 AACAGTGGCTTTGTT 15 ZNF713_C2H2_full_monomeric_17_1 SELEX + 598205 598221 6.0E-06 AATAAAAATGCCATAAA 17 ZNF713_C2H2_full_monomeric_17_1 SELEX + 598314 598330 5.0E-06 TTGAAGAAAGTCACGAT 17 SOX2_HMG_full_dimeric_17_1 SELEX + 597259 597275 1.0E-05 GAACAAAGCCACTGTTC 17 SOX2_HMG_full_dimeric_17_1 SELEX - 597259 597275 1.0E-05 GAACAGTGGCTTTGTTC 17 SOX9_HMG_full_dimeric_16_1 SELEX - 596659 596674 9.0E-06 AAGAACATTCAGTTTT 16 MEF2D_MADS_DBD_dimeric_12_1 SELEX + 598214 598225 3.0E-06 GCCATAAATAGA 12 ESR2_MA0258.1 JASPAR + 598319 598336 3.0E-06 GAAAGTCACGATGATCTG 18 POU3F4_POU_DBD_monomeric_11_1 SELEX - 598155 598165 7.0E-06 TGCATATTTTT 11 SOX8_HMG_full_dimeric_15_1 SELEX + 597260 597274 5.0E-06 AACAAAGCCACTGTT 15 Esrrb_MA0141.1 JASPAR - 598309 598320 2.0E-06 TCTTCAAGGTCA 12 MEF2A_MADS_DBD_dimeric_12_1 SELEX + 598152 598163 1.0E-05 TCTAAAAATATG 12 MEF2A_MADS_DBD_dimeric_12_1 SELEX + 598214 598225 4.0E-06 GCCATAAATAGA 12 Sox17_HMG_DBD_dimeric_15_1 SELEX - 597260 597274 8.0E-06 AACAGTGGCTTTGTT 15 Pknox2_MEIS_DBD_dimeric_12_1 SELEX + 597124 597135 8.0E-06 TGACAGGTGAGA 12 TEAD1_TEA_full_monomeric_10_1 SELEX + 596688 596697 2.0E-06 CACATTCCTT 10 RARG_nuclearreceptor_full_dimeric_18_1 SELEX - 598298 598315 0.0E+00 AAGGTCACGCAAAGTTTA 18 MEF2B_MADS_full_dimeric_12_1 SELEX + 598214 598225 3.0E-06 GCCATAAATAGA 12 Meis3_MEIS_DBD_dimeric_12_1 SELEX + 597124 597135 3.0E-06 TGACAGGTGAGA 12 ESRRG_nuclearreceptor_full_dimeric_18_1 SELEX - 598298 598315 4.0E-06 AAGGTCACGCAAAGTTTA 18 HOXC12_homeodomain_DBD_monomeric_9_1 SELEX + 598203 598211 7.0E-06 CTAATAAAA 9 MEF2A_MA0052.1 JASPAR - 598153 598162 8.0E-06 ATATTTTTAG 10 MEF2A_MA0052.1 JASPAR - 598215 598224 5.0E-06 CTATTTATGG 10 HOXD13_homeodomain_DBD_monomeric_10_1 SELEX + 598203 598212 1.0E-06 CTAATAAAAA 10 Rarb_nuclearreceptor_DBD_dimeric_17_1 SELEX - 598298 598314 0.0E+00 AGGTCACGCAAAGTTTA 17 ESR1_MA0112.2 JASPAR - 598321 598340 9.0E-06 AAAACAGATCATCGTGACTT 20 RARA_nuclearreceptor_full_dimeric_17_1 SELEX - 598298 598314 2.0E-06 AGGTCACGCAAAGTTTA 17 SP4_C2H2_full_monomeric_17_1 SELEX + 596952 596968 2.0E-06 CAGGCCCCGCCCACCCG 17 SP4_C2H2_full_monomeric_17_1 SELEX + 596976 596992 1.0E-06 CAGGCCCCGCCCACCCT 17 SOX18_HMG_full_dimeric_15_1 SELEX - 597260 597274 1.0E-06 AACAGTGGCTTTGTT 15 ESRRG_nuclearreceptor_full_monomeric_10_1 SELEX - 598308 598317 3.0E-06 TCAAGGTCAC 10 SOX15_HMG_full_dimeric_15_1 SELEX + 597260 597274 7.0E-06 AACAAAGCCACTGTT 15 SP1_C2H2_DBD_monomeric_11_1 SELEX + 596775 596785 1.0E-05 GCCCCGCCCAC 11 SP1_C2H2_DBD_monomeric_11_1 SELEX + 596826 596836 1.0E-05 GCCCCGCCCAC 11 SP1_C2H2_DBD_monomeric_11_1 SELEX + 596979 596989 1.0E-05 GCCCCGCCCAC 11 SOX7_HMG_full_dimeric_17_1 SELEX + 597259 597275 7.0E-06 GAACAAAGCCACTGTTC 17 SOX7_HMG_full_dimeric_17_1 SELEX - 597259 597275 1.0E-06 GAACAGTGGCTTTGTTC 17 HOXA13_homeodomain_full_monomeric_10_1 SELEX + 598203 598212 1.0E-06 CTAATAAAAA 10 Msx3_homeodomain_DBD_dimeric_18_1 SELEX + 598203 598220 6.0E-06 CTAATAAAAATGCCATAA 18 TEAD3_TEA_DBD_monomeric_8_1 SELEX - 598272 598279 1.0E-05 ACATTCCA 8 Esrra_nuclearreceptor_DBD_monomeric_11_1 SELEX - 598308 598318 1.0E-06 TTCAAGGTCAC 11 ESRRB_nuclearreceptor_DBD_monomeric_11_1 SELEX - 598307 598317 4.0E-06 TCAAGGTCACG 11 Meis2_MEIS_DBD_dimeric_12_1 SELEX + 597124 597135 2.0E-06 TGACAGGTGAGA 12 HOXC13_homeodomain_DBD_monomeric_10_1 SELEX + 598203 598212 1.0E-06 CTAATAAAAA 10 MEIS2_MEIS_DBD_dimeric_14_1 SELEX + 597123 597136 4.0E-06 ATGACAGGTGAGAA 14 CPEB1_RRM_full_monomeric_8_1 SELEX + 598205 598212 4.0E-06 AATAAAAA 8 TEAD4_TEA_DBD_monomeric_10_1 SELEX + 596688 596697 2.0E-06 CACATTCCTT 10 Sox3_HMG_DBD_dimeric_17_1 SELEX - 600402 600418 7.0E-06 GAACAATAGAAGACTTT 17 RORA_1_MA0071.1 JASPAR - 598309 598318 3.0E-06 TTCAAGGTCA 10 MSX1_homeodomain_DBD_dimeric_18_1 SELEX + 598203 598220 3.0E-06 CTAATAAAAATGCCATAA 18 CDX2_homeodomain_DBD_monomeric_9_1 SELEX + 598203 598211 9.0E-06 CTAATAAAA 9 HOXB13_homeodomain_DBD_monomeric_10_1 SELEX + 598203 598212 1.0E-06 CTAATAAAAA 10 SOX9_HMG_full_dimeric_17_1 SELEX + 597259 597275 6.0E-06 GAACAAAGCCACTGTTC 17 SOX9_HMG_full_dimeric_17_1 SELEX - 597259 597275 7.0E-06 GAACAGTGGCTTTGTTC 17 Zfx_MA0146.1 JASPAR - 597003 597016 4.0E-06 GCGGCCCGGGCCTG 14 V_HOXA9_01_M01351 TRANSFAC + 598211 598227 3.0E-06 AATGCCATAAATAGAGA 17 RXR_RAR_DR5_MA0159.1 JASPAR - 598298 598314 1.0E-06 AGGTCACGCAAAGTTTA 17 V_MEIS1_02_M01419 TRANSFAC - 597120 597135 6.0E-06 TCTCACCTGTCATTCC 16 V_PREP1_01_M01459 TRANSFAC - 597120 597135 3.0E-06 TCTCACCTGTCATTCC 16 V_CDX2_Q5_01_M01659 TRANSFAC + 598214 598224 5.0E-06 GCCATAAATAG 11 V_AHRARNT_01_M00235 TRANSFAC + 598298 598313 5.0E-06 TAAACTTTGCGTGACC 16 V_ESR1_01_M02261 TRANSFAC - 598321 598340 9.0E-06 AAAACAGATCATCGTGACTT 20 V_SOX30_03_M02804 TRANSFAC - 600406 600421 1.0E-05 TGAGAACAATAGAAGA 16 V_HMGIY_Q3_M01010 TRANSFAC - 598204 598218 5.0E-06 ATGGCATTTTTATTA 15 V_ERR3_Q2_01_M02094 TRANSFAC - 598307 598319 0.0E+00 CTTCAAGGTCACG 13 V_LRH1_Q5_01_M02098 TRANSFAC - 598309 598319 1.0E-06 CTTCAAGGTCA 11 V_AR_Q6_01_M01996 TRANSFAC + 596660 596674 2.0E-06 AAACTGAATGTTCTT 15 V_MEF2_02_M00231 TRANSFAC + 598208 598229 6.0E-06 AAAAATGCCATAAATAGAGAAT 22 V_ISL2_01_M01328 TRANSFAC + 598288 598303 1.0E-06 AAAAATCACTTAAACT 16 V_GKLF_02_M01588 TRANSFAC + 596775 596786 8.0E-06 GCCCCGCCCACC 12 V_GKLF_02_M01588 TRANSFAC + 596826 596837 8.0E-06 GCCCCGCCCACC 12 V_GKLF_02_M01588 TRANSFAC + 596979 596990 8.0E-06 GCCCCGCCCACC 12 V_NERF_Q2_M00531 TRANSFAC - 596687 596704 3.0E-06 CGTCAGGAAGGAATGTGG 18 V_NKX24_01_M01350 TRANSFAC + 598289 598304 4.0E-06 AAAATCACTTAAACTT 16 V_HOXD13_01_M01404 TRANSFAC + 598200 598215 2.0E-06 TACCTAATAAAAATGC 16 V_CDX_Q5_M00991 TRANSFAC + 598204 598221 3.0E-06 TAATAAAAATGCCATAAA 18 V_HOXC13_01_M01317 TRANSFAC + 598199 598214 6.0E-06 TTACCTAATAAAAATG 16 V_MEF2A_Q6_M02024 TRANSFAC - 598152 598161 0.0E+00 TATTTTTAGA 10 V_AR_03_M00956 TRANSFAC + 596653 596679 5.0E-06 TTTTGAAAAACTGAATGTTCTTTCCAC 27 V_SOX13_03_M02797 TRANSFAC - 600407 600422 6.0E-06 TTGAGAACAATAGAAG 16 V_GC_01_M00255 TRANSFAC - 596773 596786 9.0E-06 GGTGGGCGGGGCCT 14 V_GC_01_M00255 TRANSFAC - 596824 596837 9.0E-06 GGTGGGCGGGGCCT 14 V_GC_01_M00255 TRANSFAC - 596977 596990 9.0E-06 GGTGGGCGGGGCCT 14 V_SOX7_03_M02807 TRANSFAC - 600403 600424 3.0E-06 GGTTGAGAACAATAGAAGACTT 22 V_MEIS2_01_M01488 TRANSFAC - 597120 597135 4.0E-06 TCTCACCTGTCATTCC 16 V_POU3F2_01_M00463 TRANSFAC - 598153 598166 9.0E-06 CTGCATATTTTTAG 14 V_PR_Q2_M00960 TRANSFAC - 596667 596676 0.0E+00 GAAAGAACAT 10 V_TGIF_02_M01346 TRANSFAC + 597119 597135 2.0E-06 AGGAATGACAGGTGAGA 17 V_HOXB13_01_M01467 TRANSFAC + 598200 598215 0.0E+00 TACCTAATAAAAATGC 16 V_SP1_Q6_M00196 TRANSFAC - 596978 596990 3.0E-06 GGTGGGCGGGGCC 13 V_ARID3A_02_M02839 TRANSFAC + 600345 600359 6.0E-06 TTGCATATCAGGTTT 15 V_MEF2_03_M00232 TRANSFAC + 598146 598167 2.0E-06 AAAGTTTCTAAAAATATGCAGC 22 V_SZF11_01_M01109 TRANSFAC - 597040 597054 7.0E-06 ACAGGGTGTCAGCAA 15 V_NKX52_01_M01315 TRANSFAC + 598289 598305 6.0E-06 AAAATCACTTAAACTTT 17 V_OCT1_04_M00138 TRANSFAC - 598152 598174 6.0E-06 TTGTGCTGCTGCATATTTTTAGA 23 V_GR_01_M00955 TRANSFAC + 596653 596679 2.0E-06 TTTTGAAAAACTGAATGTTCTTTCCAC 27 V_ERR1_Q3_M01841 TRANSFAC + 598306 598320 0.0E+00 GCGTGACCTTGAAGA 15 V_MEF2_04_M00233 TRANSFAC + 598146 598167 2.0E-06 AAAGTTTCTAAAAATATGCAGC 22 V_MEF2_04_M00233 TRANSFAC + 598208 598229 4.0E-06 AAAAATGCCATAAATAGAGAAT 22 V_SRF_Q5_02_M01007 TRANSFAC - 598211 598229 7.0E-06 ATTCTCTATTTATGGCATT 19 V_TGIF1_01_M03111 TRANSFAC + 597119 597135 2.0E-06 AGGAATGACAGGTGAGA 17 V_CTCF_02_M01259 TRANSFAC + 597083 597102 9.0E-06 CAGACGCCTCCAGGGGGCAG 20 V_SOX11_03_M02795 TRANSFAC - 600406 600422 3.0E-06 TTGAGAACAATAGAAGA 17 V_CTCF_01_M01200 TRANSFAC + 597085 597104 1.0E-06 GACGCCTCCAGGGGGCAGCA 20 V_FOXO1_Q5_M01216 TRANSFAC - 598333 598341 1.0E-05 AAAAACAGA 9 V_HOXD9_Q2_M01834 TRANSFAC + 600273 600282 8.0E-06 ACACATAAAT 10 V_SF1_Q6_01_M01132 TRANSFAC + 598309 598317 3.0E-06 TGACCTTGA 9 V_GR_Q6_02_M01836 TRANSFAC + 596663 596675 6.0E-06 CTGAATGTTCTTT 13 V_SP4_Q5_M01273 TRANSFAC + 596775 596785 9.0E-06 GCCCCGCCCAC 11 V_SP4_Q5_M01273 TRANSFAC + 596826 596836 9.0E-06 GCCCCGCCCAC 11 V_SP4_Q5_M01273 TRANSFAC + 596979 596989 9.0E-06 GCCCCGCCCAC 11 V_SOX15_03_M02799 TRANSFAC - 600406 600422 1.0E-06 TTGAGAACAATAGAAGA 17 V_MTF1_06_M02882 TRANSFAC + 598201 598214 1.0E-05 ACCTAATAAAAATG 14 V_AP3_Q6_M00690 TRANSFAC - 598407 598414 5.0E-06 TCTAAATT 8 V_MRG2_01_M01395 TRANSFAC - 597120 597135 6.0E-06 TCTCACCTGTCATTCC 16 V_HOXB8_01_M01451 TRANSFAC + 598210 598225 4.0E-06 AAATGCCATAAATAGA 16 V_SMAD4_04_M02898 TRANSFAC - 598130 598146 5.0E-06 TTCCACCCGCCATTGTC 17 V_HDX_01_M01333 TRANSFAC + 598284 598300 1.0E-06 CAGTAAAAATCACTTAA 17 V_ERR1_Q2_M00511 TRANSFAC - 598307 598320 8.0E-06 TCTTCAAGGTCACG 14 V_ERR2_01_M01589 TRANSFAC - 598306 598317 0.0E+00 TCAAGGTCACGC 12 V_T3RBETA_Q6_01_M02119 TRANSFAC - 597053 597069 5.0E-06 AAGTCAGCCGAGGGCAC 17 V_ESRRA_03_M02748 TRANSFAC - 598304 598320 0.0E+00 TCTTCAAGGTCACGCAA 17 V_SOX21_04_M02907 TRANSFAC + 600407 600423 8.0E-06 CTTCTATTGTTCTCAAC 17 V_RXR_RAR_01_M02272 TRANSFAC - 598298 598314 1.0E-06 AGGTCACGCAAAGTTTA 17 V_SOX8_03_M02808 TRANSFAC + 600406 600422 1.0E-06 TCTTCTATTGTTCTCAA 17 V_ERR1_Q2_01_M02093 TRANSFAC - 598306 598316 1.0E-06 CAAGGTCACGC 11 V_SOX18_04_M02905 TRANSFAC + 598401 598416 7.0E-06 GGTCTGAATTTAGACG 16 V_PR_02_M00957 TRANSFAC + 596653 596679 4.0E-06 TTTTGAAAAACTGAATGTTCTTTCCAC 27 V_PPARG_02_M00515 TRANSFAC - 598317 598339 8.0E-06 AAACAGATCATCGTGACTTTCTT 23 V_BARHL2_01_M01446 TRANSFAC + 598288 598303 6.0E-06 AAAAATCACTTAAACT 16 V_SRY_05_M02917 TRANSFAC - 600407 600423 6.0E-06 GTTGAGAACAATAGAAG 17 V_HMX3_02_M01413 TRANSFAC + 598289 598305 8.0E-06 AAAATCACTTAAACTTT 17 V_MAFK_04_M02880 TRANSFAC + 598205 598219 9.0E-06 AATAAAAATGCCATA 15 V_SP1_Q4_01_M00932 TRANSFAC - 596978 596990 2.0E-06 GGTGGGCGGGGCC 13 V_TCF3_05_M02920 TRANSFAC + 598200 598214 7.0E-06 TACCTAATAAAAATG 15 V_AR_Q2_M00447 TRANSFAC + 596659 596673 2.0E-06 AAAACTGAATGTTCT 15 V_ESR2_01_M02377 TRANSFAC + 598319 598336 3.0E-06 GAAAGTCACGATGATCTG 18 V_NANOG_02_M01247 TRANSFAC - 600405 600424 9.0E-06 GGTTGAGAACAATAGAAGAC 20

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