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HPSE Entrez Description Chrom. Strand Promoter (Start - Stop) TSS 10855 heparanase chr4 - 84255466 - 84261306 84255966 Transcription Factor Binding Sites   Download all TFBS in the HPSE promoter   Motif Source Strand Start Stop PValue Match Sequence Overlap w/ Footprints NKX2-8_homeodomain_full_monomeric_9_1 SELEX + 84256215 84256223 3.0E-06 CCACTTGAA 9 Egr1_MA0162.1 JASPAR - 84253306 84253316 9.0E-06 TGTGTGGGCGG 11 SP3_C2H2_DBD_monomeric_11_1 SELEX + 84256208 84256218 6.0E-06 CCCACACCCAC 11 EWSR1-FLI1_MA0149.1 JASPAR - 84256156 84256173 2.0E-06 GAAAGCGAGCAAGGAAGT 18 ZNF238_C2H2_full_monomeric_13_1 SELEX + 84251065 84251077 3.0E-06 GAGCCAGATGTGG 13 NR2E1_nuclearreceptor_full_dimeric_14_1 SELEX + 84251261 84251274 1.0E-05 AAGGTAACAAGTCA 14 Lhx3_MA0135.1 JASPAR - 84253338 84253350 8.0E-06 TGTTTAATTTATT 13 Lhx3_MA0135.1 JASPAR + 84253341 84253353 8.0E-06 AAATTAAACAACA 13 EGR4_C2H2_DBD_monomeric_16_2 SELEX + 84253304 84253319 6.0E-06 TCCCGCCCACACATCT 16 ZNF143_C2H2_DBD_monomeric_16_1 SELEX + 84251108 84251123 1.0E-05 CACCCGTAATGCACAT 16 NKX2-3_homeodomain_full_monomeric_10_1 SELEX + 84256214 84256223 8.0E-06 CCCACTTGAA 10 PRDM1_C2H2_full_monomeric-or-dimeric_15_1 SELEX - 84256187 84256201 1.0E-05 CGTAAGTGAACGTGA 15 FOXC1_MA0032.1 JASPAR + 84251018 84251025 7.0E-06 AGTAAGTA 8 ELF5_MA0136.1 JASPAR + 84256155 84256163 4.0E-06 TACTTCCTT 9 HOXA10_homeodomain_DBD_monomeric_11_1 SELEX + 84253335 84253345 8.0E-06 TGTAATAAATT 11 V_HOXA9_01_M01351 TRANSFAC + 84253333 84253349 8.0E-06 ACTGTAATAAATTAAAC 17 V_TST1_02_M01316 TRANSFAC + 84253334 84253350 9.0E-06 CTGTAATAAATTAAACA 17 V_DBX1_01_M01483 TRANSFAC + 84253337 84253353 1.0E-06 TAATAAATTAAACAACA 17 V_MAX_Q6_M01830 TRANSFAC - 84256331 84256342 6.0E-06 CAGAACACGTGC 12 V_BCL6B_03_M02740 TRANSFAC + 84253322 84253337 3.0E-06 ACTTTTGAGGAACTGT 16 V_HP1SITEFACTOR_Q6_M00725 TRANSFAC - 84253334 84253345 4.0E-06 AATTTATTACAG 12 V_ETS_B_M00340 TRANSFAC - 84255995 84256008 7.0E-06 GGGAGGAAGTGCTA 14 V_SP1_Q6_01_M00931 TRANSFAC - 84256127 84256136 7.0E-06 GGGGCGGGGT 10 V_NFMUE1_Q6_M00651 TRANSFAC - 84256232 84256240 6.0E-06 CGGCCATCT 9 V_RP58_01_M00532 TRANSFAC - 84251067 84251078 5.0E-06 TCCACATCTGGC 12 V_NKX25_Q6_M02108 TRANSFAC + 84256214 84256224 1.0E-06 CCCACTTGAAG 11 V_EGR1_02_M01972 TRANSFAC - 84253306 84253316 7.0E-06 TGTGTGGGCGG 11 V_LHX3_01_M01471 TRANSFAC + 84253336 84253352 4.0E-06 GTAATAAATTAAACAAC 17 V_AIRE_02_M01000 TRANSFAC - 84251256 84251280 3.0E-06 CCCTTTTGACTTGTTACCTTGGGCA 25 V_ZFP691_03_M02833 TRANSFAC - 84251047 84251063 5.0E-06 CATCCAGTGCTCCTGCT 17 V_PAX6_02_M01391 TRANSFAC - 84253335 84253350 2.0E-06 TGTTTAATTTATTACA 16 V_PAX6_02_M01391 TRANSFAC + 84253337 84253352 8.0E-06 TAATAAATTAAACAAC 16 V_HNF1_C_M00206 TRANSFAC - 84253337 84253353 5.0E-06 TGTTGTTTAATTTATTA 17 V_AP1_Q6_M00174 TRANSFAC - 84251093 84251103 5.0E-06 CATGACTCAGT 11 V_RPC155_01_M01798 TRANSFAC - 84251279 84251294 7.0E-06 GCTATAGTTCAAGTCC 16 V_ETS_Q4_M00771 TRANSFAC + 84255995 84256006 1.0E-05 TAGCACTTCCTC 12 V_BACH1_01_M00495 TRANSFAC + 84251091 84251105 6.0E-06 ACACTGAGTCATGGC 15 V_LMX1_01_M01409 TRANSFAC + 84253336 84253352 6.0E-06 GTAATAAATTAAACAAC 17 V_STAF_01_M00262 TRANSFAC + 84251106 84251127 6.0E-06 ATCACCCGTAATGCACATCAGG 22 V_HNF1_Q6_M00790 TRANSFAC - 84253333 84253350 1.0E-06 TGTTTAATTTATTACAGT 18 V_KROX_Q6_M00982 TRANSFAC + 84253305 84253318 6.0E-06 CCCGCCCACACATC 14 V_AP1_Q6_01_M00925 TRANSFAC - 84251094 84251102 7.0E-06 ATGACTCAG 9 V_SP1_Q6_M00196 TRANSFAC - 84256126 84256138 9.0E-06 GCGGGGCGGGGTT 13 V_CDX2_01_M01449 TRANSFAC + 84253332 84253347 2.0E-06 AACTGTAATAAATTAA 16 V_MAZ_Q6_01_M02023 TRANSFAC - 84256031 84256044 2.0E-06 GGCGGAGGAAAGGA 14 V_MAZ_Q6_01_M02023 TRANSFAC - 84256078 84256091 4.0E-06 GTGGGAGGGATGGA 14 V_HMX1_01_M00433 TRANSFAC - 84256212 84256221 5.0E-06 CAAGTGGGTG 10 V_HNF1_Q6_01_M01011 TRANSFAC + 84253332 84253352 2.0E-06 AACTGTAATAAATTAAACAAC 21 V_CDX1_01_M01373 TRANSFAC + 84253332 84253347 3.0E-06 AACTGTAATAAATTAA 16 V_AP1_C_M00199 TRANSFAC - 84251094 84251102 6.0E-06 ATGACTCAG 9 V_NCX_02_M01420 TRANSFAC + 84253337 84253353 3.0E-06 TAATAAATTAAACAACA 17 V_HB24_01_M01399 TRANSFAC + 84253334 84253348 1.0E-06 CTGTAATAAATTAAA 15 V_HB24_01_M01399 TRANSFAC - 84253335 84253349 2.0E-06 GTTTAATTTATTACA 15 V_POU6F1_01_M00465 TRANSFAC - 84253339 84253349 5.0E-06 GTTTAATTTAT 11 V_SP1_01_M00008 TRANSFAC - 84256127 84256136 3.0E-06 GGGGCGGGGT 10 V_EGR2_01_M00246 TRANSFAC - 84253306 84253317 8.0E-06 ATGTGTGGGCGG 12 V_AP1_Q2_01_M00924 TRANSFAC - 84251090 84251101 5.0E-06 TGACTCAGTGTG 12 V_ELF1_Q6_M00746 TRANSFAC + 84251078 84251089 9.0E-06 AGATGAGGAAAA 12 V_AP1_Q4_01_M00926 TRANSFAC + 84251095 84251102 1.0E-05 TGAGTCAT 8 V_TAL1_01_M01591 TRANSFAC - 84251076 84251088 4.0E-06 TTTCCTCATCTCC 13 V_BRN4_01_M01473 TRANSFAC - 84253333 84253349 5.0E-06 GTTTAATTTATTACAGT 17 V_HOXB8_01_M01451 TRANSFAC + 84253332 84253347 1.0E-06 AACTGTAATAAATTAA 16 V_FOXL1_04_M02753 TRANSFAC + 84253339 84253355 7.0E-06 ATAAATTAAACAACAGG 17 V_BDP1_01_M01796 TRANSFAC + 84251279 84251290 2.0E-06 GGACTTGAACTA 12 V_LMX1B_01_M01363 TRANSFAC - 84253336 84253352 4.0E-06 GTTGTTTAATTTATTAC 17 V_FRA1_Q5_M01267 TRANSFAC + 84251095 84251102 1.0E-05 TGAGTCAT 8 V_FPM315_01_M01587 TRANSFAC + 84256277 84256288 9.0E-06 CGGGGAGGAGCG 12 V_T3RBETA_Q6_01_M02119 TRANSFAC + 84251021 84251037 9.0E-06 AAGTAAGAGGGGGACAG 17 V_T3RBETA_Q6_01_M02119 TRANSFAC + 84255659 84255675 4.0E-06 AGGTGTCAGGAGGACAG 17 V_ETS1_B_M00339 TRANSFAC - 84255993 84256007 5.0E-06 GGAGGAAGTGCTAGA 15 V_EWSR1FLI1_01_M02252 TRANSFAC - 84256156 84256173 2.0E-06 GAAAGCGAGCAAGGAAGT 18 V_SOX18_04_M02905 TRANSFAC - 84253336 84253351 7.0E-06 TTGTTTAATTTATTAC 16 V_EKLF_Q5_M01874 TRANSFAC + 84256209 84256218 9.0E-06 CCACACCCAC 10 V_NFE2_01_M00037 TRANSFAC + 84251092 84251102 5.0E-06 CACTGAGTCAT 11 V_ELF5_04_M02241 TRANSFAC + 84256155 84256163 4.0E-06 TACTTCCTT 9 V_LHX5_01_M01353 TRANSFAC + 84253336 84253352 4.0E-06 GTAATAAATTAAACAAC 17 V_HOXA10_01_M01464 TRANSFAC + 84253333 84253348 0.0E+00 ACTGTAATAAATTAAA 16 V_HOXD10_01_M01375 TRANSFAC + 84253333 84253349 9.0E-06 ACTGTAATAAATTAAAC 17 V_BRF1_01_M01747 TRANSFAC - 84251278 84251290 5.0E-06 TAGTTCAAGTCCC 13 V_HOXC9_01_M01416 TRANSFAC + 84253332 84253347 5.0E-06 AACTGTAATAAATTAA 16

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